Ed and Al become interested in biological transmutation after witnessing a chimera in Lior, so Colonel Mustang introduces them to an expert in the field, Shou Tucker, the Life-binding Alchemist. Meanwhile, a serial killer whose targets are only State Alchemists is on the loose in Central.

After learning about what happened to Shou Tucker and his daughter Nina, the Elric brothers begin to question what alchemy really is. Meanwhile, Colonel Mustang and the other State Alchemists track down the serial killer Scar.

Ed's journey to a distant land of legend offers surprising new details behind Mustang's alleyway encounter with Ross. After a fateful encounter of his own, Elric learns a stunning secret about Winry's parents.

Mustang, Alphonse, Havoc, and Hawkeye split up to search a military laboratory. If Lust has her way, none of them will ever see each other alive again.

Hoping to learn more about the Homunculi, Ed and Al devise a risky strategy to lure the creatures out of hiding. Step one of the brothers' plan: risk death at the hands of their most dangerous enemy - Scar.

The Homunculi are winning. Ed and Ling face a horrific enemy within the hellish expanse of Gluttony's gut. In Central, Mustang can only watch as his loyal band of soldiers is divided by Bradley.

Even as his subordinates are scattered, Mustang finds he still has allies - unfortunately, so do his enemies. The Homunculi unleash one of the Ishvalan War's most notorious killers to deal with Scar and Marcoh.

May and Marcoh's efforts to decode the research of Scar's brother yield progress, but it might be too late to prevent catastrophe. Violence is spreading across the land, and it will soon reach the northern border.

Those who have prepared to reclaim a nation find their plans exposed. The chaos multiplies after an assassin strikes down a mighty leader in broad daylight, and Alphonse falls prey to monsters in the dark.

After investigating the twisted roots of the Bradley family tree, Mustang gathers his most loyal troops. Meanwhile, Ed and his allies continue fighting Selim, who shocks everyone by turning on his own kind.

Mustang leads a ragtag assault force of hardy northern troops - and welcome faces from the past - against Central's defenses. Victory seems certain until the military's most horrifying secret is revealed.

Ed's crew faces a horde of ravenous mannequins, abominations powered by human souls. The twisted creatures aren't the only unstoppable force on the battlefield -Pride, Sloth, and Envy are on the move.

After a startling radio broadcast grants Mustang's forces the advantage over Central's troops, the Flame Alchemist unleashes the brutal power of burning hatred in the battle against the killer of Maes Hughes.

Central's forces fight with renewed vigor following Bradley's return, and the Fuhrer himself enters the battle. His first opponent is an armored tank. His second foe is even tougher.

As one shared journey ends, many journeys begin. Those most touched by the conflict which surrounded the Philosopher's Stone now go their separate ways. One thing is certain: they will meet again.