Luffy faces against Urashima in a sumo match. Holdem tries to pull off Tama's cheek until Luffy sends Urashima crashing into Holdem's residence.

Luffy, Zoro, and Kiku fight back against the attacking Beasts Pirates and their samurai allies. Luffy calls out to Holdem, who approaches Luffy's group while holding Tama hostage.

Holdem continues to taunt Luffy's group while holding Tama hostage, threatening to chew her up with Kamijiro. Kiku warns Luffy against attacking Holdem as Kuri is ruled by Jack, and he will destroy Bakura Town if they fight back. Speed arrived with the Treasure ship of food from Paradise Farm. Luffy ordered Zoro to steal the food cart, which he did with Kiku's help. Luffy proceed to punch Kamijiro and rescue Tama. When he saw the injuries on Tama's cheek from being pulled by Holdem with a pincer, he flew into a rage and punched Holdem with Red Hawk, where Tama see Ace in Luffy.

After defeating Holdem, Luffy grabs Tama and flees the scene. During the escape, they encounter Speed and Tama tames her. They then catch up to Zoro, Kiku, and Komachiyo as the latter three were fleeing with the stolen food cart. Meanwhile, Law fights Hawkins but their battle is interrupted when the food cart passes by them. After jumping on the food cart, Law reprimands Zoro for his group's actions.

This is a story of Luffy in an alternate reality. While sailing on the sea by himself, a strange bird crashes into his small boat. He looks up to find a giant ship where a girl named Ann has been captured. It seems like the bird belongs to her.

Luffy's group delivers the stolen food cart to Okobore Town and Luffy also gives the residents clean water. After Luffy parted ways with Tama and Speed, Law takes the group to the ruins of Oden Castle.

While traveling to Oden Castle, Komachiyo, Law, Luffy, Zoro, and Kiku encounter a ferocious tiger. Zoro separates from the group to fight the tiger. The rest of the group later arrive at the ruins of Oden Castle. After seeing the graves of the Kozuki Family, Luffy reunites with Kin'emon, Momonosuke, Sanji, Nami, Brook, Chopper, and Carrot. Once they are inside Oden Castle, Kin'emon reveals that he, Momonosuke, Kiku, Raizo, and Kanjuro came from the past.

Kin'emon explains to Luffy's group about the past. He recounts stories such as how his Lord, Kozuki Oden, brings order to Kuri, the day of his demise, and Toki sending Kin'emon's group and Momonosuke to the future.

Kin'emon explained to Luffy's group, who just arrived, the preparation they have done till now and assigned new tasks to them. He laid out the plan of how they are going to defeat Kaido, but there are still lots of work to do such as recruiting more allies and gathering more information on the enemy.

While Usopp is looking for allies in Wano Country, Shutenmaru and the Mt. Atama Thieves plunder Okobore Town. Jack suddenly arrives at Okubore Town and confronts Shutenmaru. Shutenmaru and Jack quickly get into a fight, but are interrupted when Kaido flies in his dragon form.

The drunken Kaido flies above Okobore Town in his dragon form and threatens Shutenmaru and all the people in the town. Luffy, Law, Kin'emon, and Kiku start running towards the town to rescue them. However, Kaido heads towards the ruins of Oden Castle and destroys it with a single attack.

Right after the destruction of Oden Castle, Luffy enters a furious battle with Kaido.

Luffy continues his furious battle with Kaido. Despite his best efforts, the battle ends in Luffy's defeat.

After his defeat, Luffy is sent to a prison in the Udon region. Kiku returns to the mountain where Oden Castle was destroyed and discovers that the group there managed to escape the destruction. Franky, Robin, Usopp, and Zoro read the news about Luffy's fight with Kaido. Inuarashi and his group discover Tama alive and take her into the forest. At the prison in Udon, Luffy meets Kid again.

Perona learns from a newspaper that Gecko Moria is alive and prepares to leave Kuraigana Island to search for him. Moria and his zombie army attack Pirate Island to find Absalom only to learn that he had been killed.

Dogstorm clashes with Shutenmaru, who refuses to aid in Kin'emon's operation. Sanji and the others seek out recruits.

Nami and Shinobu finish with their current objective in the Flower Capital and Sanji's soba stand is growing more popular. In Udon, Luffy and Kid continue to work. Dobon becomes angry at them for eating alot of food. The vice warden tries to attack them only to be defeated by the pair.

People of the Land of Wano lining up just to taste the delicious soba made by Sanji. He decided to sell the noodles to attract samurais who could join them in their upcoming war against Shogun Kurozumi Orochi, Emperor Kaido, and the Beast Pirates. While he was serving a little girl, three men arrived at his soba stand, they urged Sanji to pay them for doing business in their territory which he refused. Sanji got into a fight when the three men ruined his soba stand.

Wano's most popular courtesan heads to the palace, but a revenge plot threatens the proceedings.

Zoro goes to a gambling house with Tonoyasu. After a brawl, they go to Ebisu Town. Meanwhile, the Ninja-Pirate-Mink-Samurai Alliance works to steal weapons and food. Franky learns that Minatomo no longer has the blueprints of Kaido's mansion. He goes all over the Flower Capital searching for them but hits a dead end. At the shogun's castle, CP-0 attempts to negotiate with Orochi.

The banquet at Orochi's palace begins and Robin begins her mission. Shinobu, Nami, and Brook sneak into the palace as well. Meanwhile, in Udon prison, Kid explains to Luffy and Caribou about how he lost his arm. Back at the Flower Capital, two of the Flying Six, Page One and X Drake arrive to hunt down Sanji.

In the seas surrounding Wano Country, the Big Mom Pirates draw closer.

As the Big Mom Pirates approach the Land of Wano, Kaido’s subordinates led by King and Queen try to get rid of them. Meanwhile, in the Flower Capital, another member of the Animal Kingdom Pirates, Page One, attacks every soba restaurant in the city, in search of Sanji.

As he confronts Page One, Sanji puts on his Raid Suit and displays its capabilities.

Back in the shogun's caste, Robin was sneaking around trying to find important information but she is soon confronted by the Orochi Oniwabanshu.

Robin gets caught by the Orochi Oniwabanshu in Orochi's castle. The ninjas attack Robin in order to find the truth of her intentions but with the help of her Devil Fruit, she is able to get away from them. Meanwhile, Chopper, Kiku, Tama, and Momonosuke find Big Mom who gets drifted to the shore after the encounter with King. Chopper and the others then discover that Big Mom lost her memories.