Jane is finally in a rhythm with Mateo, but finds herself out of step with Rafael and Michael and the potential love triangle that is brewing between them. In order to not be sued for missing their Vegas arrangement, Xo and Rogelio must perform on a cruise ship. Petra and Rafael are suspicious of Lachlan and his recent business proposal, but are soon faced with an even more shocking revelation. Meanwhile, Alba makes a very big decision that will impact her life.

As Jane tries to juggle motherhood and school, she forgets to plan Lina's surprise 25th birthday party leading to friction between the best friends. Rafael tells Jane that he still wants a romantic relationship with her, but she is blindsided when he asks for joint custody. Petra tries to enlist Michael's help to keep Jane and Rafael apart. Rogelio's ex-wife, Luciana, is blackmailing him with secrets from years ago to get her role back on "The Passions of Santos." Meanwhile, Michael gets a new partner and a possible lead in the Sin Rostro case.

Jane and Rafael begin estate planning for Mateo, but when Jane realizes that Mateo will inherit millions, she worries he will become spoiled. Jane is also finally starting graduate school, but her professor is not sympathetic to her situation. Rogelio's famous show is ending, so he decides to create his own show, with Xo's encouragement. Petra is shocked to learn what she is having and turns to Jane for advice and help in getting her mother out of prison. Meanwhile, after Michael's issues at work, he has decided to go rogue and is even more determined to catch Sin Rostro.

Jane and Rafael are still at odds with each other as they are about to celebrate Mateo's first Christmas. Jane discovers that the scholarship she thought she had been awarded isn't quite that and now needs to find a way to pay for grad school. Rogelio has employed an intern and is having trouble trusting him. Petra is struggling with the latest dilemma her mother has gotten them into. Meanwhile, Michael is trying to find Luisa's mother who he now suspects is not dead and may be involved with Sin Rostro.

In the middle of the night, Jane gets a burst of creative energy and starts her thesis. Unfortunately, Mateo spills orange juice all over her computer causing it to crash and Jane loses all her hard work. When Jane brings her computer to be fixed, she meets Dax and with the urging of her friend Lina (guest star Diane Guerrero), Jane considers dating again. Rogelio decides to have his mother Liliana become his manager, but soon realizes that was not the best decision. When Rafael comes to after being attacked by his mother now known to be the crime lord Mutter, he goes to Michael.

Jane is considering whether or not to take her relationship with Jonathan to the next level and turns to Xo for advice. Xo and Rogelio have their own relationship issues to work out. Meanwhile, as a favor to Rafael, Jane agrees to help Petra by throwing her a baby shower. Luisa is in a terrible car accident, but all is not as it seems. Michael comes to a realization he must share with Jane.

Jane and Michael are back together and Jane wishes she had never left Michael in the first place. Rafael appears to have regressed to his playboy ways and Petra is concerned about him. But Petra is not aware that Rafael is helping Michael find Mutter's other son, Derek. Rogelio's new assistant, Paolo, is actually his stalker Lola. Finally, Petra gives birth to the twins.

Jane and Michael's wedding plans are side tracked when the Villanueva house is ruined by a flood. Jane is looking to have a small wedding, but Rogelio wants nothing more than to throw Jane a big blow out wedding so he can invite all his celebrity friends. Jane and Michael are also looking for the perfect home to move into and when they find a place 45 minutes away, Rafael has an issue being that far from Mateo. Petra is still having trouble adjusting to motherhood.

With Jane and Xo still dealing with the fallout from Jane's bachelorette party, Jane is now left with finding someone to watch Mateo. When Michael volunteers to help everything is perfect until Mateo does something for the first time in front of Rafael, upsetting him. Rogelio gets in trouble with the network executives when he uses the crew for his own use, so he turns to his favorite writer Dina to help fix the problem. Meanwhile, someone is causing problems for Petra at the Marbella and in her personal life. 

The Villanueva women have a tradition for Mother's Day and Jane is excited to get to participate in her first one as a mom. However, their plans are thwarted when Petra invites everyone to a brunch instead. Jane's TA position is put into jeopardy when a mysterious ad appears in the college newspaper. Rogelio is hurt when Dina doesn't invite him to her birthday party, so he asks Jane to help in an unusual way. Rafael finds himself in big trouble thanks to his half-brother. Meanwhile, Xo finds herself in a compromising position.

After moving up the wedding day, Jane and Michael find all their carefully made plans thrown into chaos. To make matters worse, Jane is unhappy with her current graduate thesis, but changing it might put a wrinkle in her big day. Rafael still has feelings for Jane, but is torn about whether he should speak now or forever hold his peace. Petra has her own family drama when Anezka has a medical emergency and ends up in the hospital. Meanwhile, Rogelio and Xo are finally coming to terms with the state of their relationship until Xo gets surprising news.

When Rafael finally admits that he is over Jane, their co-parenting style is tested when they argue over what the best preschool is for Mateo. Xo is paranoid about Alba finding out her secret and what it will do to their relationship. Rogelio is ready to cross over to American TV, especially after he learns his nemesis is making the transition. Meanwhile, “Anezka” is trying to find new dirt on Rafael when she begins an unexpected romance.

Jane and Michael’s housewarming party in their new home hits a snag when they discover they are being evicted for not paying their rent. Rogelio is still trying to break into American television, but needs some unusual help from Rafael. Xo makes a new career choice, but Alba and Jane aren’t convinced this is the one for her. Meanwhile, “Petra” shocks Rafael with what she wants to do with her shares of the hotel.

When Jane’s cousin unexpectedly shows up at her door, Jane questions her own less than spontaneous life. Xo finds the perfect location for her dance studio, but it happens to be near her ex-boyfriend, Bruce. Rogelio realizes his feelings for Xo, but struggles with telling her. Petra is served with a lawsuit and she decides to ask Rafael for help. Meanwhile, Michael is ready to get to work just as they get a break in the Mutter case.

Jane finds it difficult to stop meddling in other people’s relationships, especially Xo’s and Rafael’s. Rogelio makes an unorthodox agreement with his matchmaker. Petra goes to great lengths to find out if Rafael is working against her. Michael, with Jane’s help, makes an important life decision with Jane’s help. Meanwhile, Rafael gets some upsetting news that leaves him reeling.

When Jane realizes that she and Michael are both stressed out, they decide to recreate their first date to help them relax. Rogelio makes a big scene on the red carpet of his film premiere and his rant goes viral, which leave him to make a choice about the direction of his career. Rafael wants to set a good example for his children leaving him to make a big decision with the ultimate sacrifice. Meanwhile, Bruce’s daughter finds herself in trouble leaving Xo and Bruce’s to discuss the future of their relationship.

When Jane is offered a book deal, she thinks her prayers have been answered and debates quitting her awful publishing job. After suffering from panic attacks, Jane is ready to get to the root of the problem. Rogelio and Xo are working on fixing their friendship until Xo finds out that Rogelio has been lying to everyone all this time. Petra is running damage control on the Marbella’s reputation after a grisly discovery is found on the property. Meanwhile, Rafael is ready to be involved again in the Mirabella and his first order of business is finding the perfect person to manage the hotel lounge.

Jane feels guilty that she isn’t around for Mateo as much as she would like, so she decides to run for Room Mom against none other than perfect mom, Petra. With the help of Jane and Xo, Alba tries to impress her crush. Rafael tries to be there for both Jane and Petra. Meanwhile, Rogelio tries to slyly get information from Mateo about Xo and Bruce’s relationship.

Jane's burgeoning relationship with Adam helps bring out her fun side and reminds her that she isn't just a mom. However, when Adam gets a job opportunity, he asks for Jane's opinion on if he should stay or go. Rogelio is facing issues with his co-star at work and things are about to get even worse for him. Xo is helping to take care of Darci to prove that Rogelio is committed to being a father to the baby. Meanwhile, Petra and Rafael are working together to try and buy back the hotel, but are running into trouble, causing tension between them.

When Jane learns that Rafael introduced Mateo to his new girlfriend, Katherine, without her consent she gets angry since she thinks Katherine has a negative influence on both Rafael and Mateo. Jane turns to Petra for help with the situation with Rafael, but Petra makes Jane an offer she can't refuse. Now that the baby is born, Rogelio and Darci struggle to choose a name for the baby they both agree on. Meanwhile, Xo is focused on growing the dance studio and decides to go into business with an unlikely foe.

Jane's book is finally being released but her plans to have a party at the local bookshop where she was inspired to be a writer hit a snag. Always wanting to help Jane, Rogelio devises a plan to save the day, but one of his archrivals threatens to ruin the day. Rafael finds a job, but it is not exactly what he was hoping for. Meanwhile, Petra has come up with an interesting strategy to get the Marbella back, but Rafael wants nothing to do with it.

After Jane and Rafael decide to send Mateo to a new school, they become guilt-ridden when they are forced to lie to get him in. After Xo convinces Rogelio to attend therapy they both are surprised by their emotions. Meanwhile, Petra finds herself in big trouble with no one to turn to for help, not even Rafael.

Jane and Rafael try to be friends which works great until Jane goes on a date. Petra and Jane's new partnership is off to a rocky start, but they both know they have to find a way to work together somehow. Rogelio is convinced he has Male Post-Partum Depression, which puts him at odds with with celebrity Post-Partum Advocate River Fields. Meanwhile, Alba finally tells Jane why she turned down the marriage proposal, leaving Jane to help Alba in the romance department.

Jane's fixation on a negative review has left her with writer's block, so Rogelio suggests she take a class to help. With the encouragement of Jane, Rafael is ready to dig into his past. Rogelio is trying to make things less about him and more about Xo, but he grows suspicious when he learns she is keeping a secret from him. Meanwhile, Petra tests the loyalty of her lawyer.

Xo struggles to make an important decision and looks to Jane for guidance leaving Rogelio feeling like an outsider. Alba is frustrated with Rogelio when it seems he is once again making his career a priority over Xo. Meanwhile, Petra turns to Rafael for advice on her love life.

Jane and Rafael find themselves in a not ideal living situation in order to save money. When Rafael sees Alba try to punish Mateo, an unexpected rift is formed. Xo tries to stay positive during treatment and even makes a new friend, but things get a little too real for her and Rogelio. River and Rogelio once again are at odds over their ideas for the show, but an unlikely person helps River see the show in a different light. Meanwhile, Petra feels unsupported by JR when she questions what really happened the night Anezka died.