The fallout on the Hill from Carrick's switch to the Republican party exiles Josh deep into a White House doghouse, so C.J. gives Donna some tips on how she can support him; Jed gets a harsh reality check from C.J. when he lingers too long in a small Oklahoma town devastated by a deadly tornado; in Jed's absence, Leo juggles an impending war between Albania and Greece, a visit from the German chancellor, a tussle over tax cuts, and an over-reaching Secretary of Defense; Josh and Toby are less than thrilled when Leo brings Angela on board to assume some of Josh's responsibilities.

When a conservative Republican Congresswoman targets Ellie's research project in an effort to discredit Jed, Toby discovers that the source of the leak is uncomfortably close to home, and Will realizes that they've all greatly underestimated his boss; Josh investigates the wisdom of a recess appointment as he tries to convince an old friend to be patient with the Sixth Circuit judicial nomination process and keep his hat in the ring; C.J. completely misreads Ben's intentions; Donna tracks down an AWOL Ryan; an opening is created on the Supreme Court; in an effort to redefine Abbey's image with the press, C.J. sets her up to do a public service announcement with the Muppets, which inspires Ellie to stand up before the press in defense of scientific research unhampered by politics.