Layton uses his new position as train detective to investigate the murder while gathering intel for the revolution on the side. Meanwhile, Melanie faces a resource crisis, with potentially drastic consequences for the entire train.

Layton descends into the black market with Till, searching for both the killer and a valuable commodity for his revolution. Melanie stages a prize fight to distract the passengers from mounting class tension.

Hiding out with help from his allies, Layton lays track for revolution and an engineering emergency threatens every soul on Snowpiercer.

On a divided Snowpiercer, Melanie intensifies her search for Layton while Layton is weaponizing her secret. Third Class faces a reckoning when he presents them with a choice.

The rebel forces are on their heels when a dangerous foe joins them. A plan is hatched to take the train, but it may destroy them all.

In the revolution's aftermath, Layton and Melanie realize the greatest threat to their survival is right over their shoulder.