The Gokaigers must help Satoru Akashi search for a Precious. However, both the Zangyack Empire and a revived Ryuuwon get in their way

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With the next fortune being "Uao", the Gokaigers seek out their next clue until they find a sign at an amusement park that says "Greater Power found here", and they find Momo Maruo, the former OhPink of the Ohrangers. However, Basco has also managed to find this Greater Power, and has been approached by Goro Hoshino, the former OhRed.

In order to defeat the more powerful Action Commander Shieldon, who boasts an impenetrable shield, Doc works on developing a new more powerful weapon.

With Navi's next clue being "Moroboshi High School", Gai realizes that their next clue to the Greatest Treasure in the Universe has to do with the Megarangers. The Gokaigers visit the high school, and discover that Kenta Date, the former Mega Red, has become a teacher, and he tells them that they must become his students in order to obtain his team's greater power

The former Time Yellow Domon tells the Gokaigers that they must go back in time to help the Goseigers and Shinkengers.

On the Gigant Horse, wondering what their next course should be, Damarsu and Isarn receive an unexpected visit from the Zangyack Imperial Fleet as the Zangyack Emperor Akudosu Gill arrives to the bridge. As news of his son's death has reached him, Akudosu has decided to assume command while having his Imperial Guards take Damarasu to the brig for failing in his mission as he then sends the Imperial Guard Zatsurigu to avenge his son.

With all 35 of the greater powers, the Gokaigers seek out the Greatest Treasure in the Universe, but Insarn is sent to begin the Zangyack Empire's final assault on the Earth

Zangyack is advancing on their final plans, which includes an all out attack on the Gokaigers, and the ultimate destruction of Earth! It has been revealed that the Greatest Treasure in the Universe can, and must, be used to wipe out Zangyack's Great Fleet, completely. The Gokaigers are hesitant to do so, as the cost of the power would be the existence of the 34 senior Super Sentai...

With their mecha defeated and the Galleon badly damaged, the Gokaigers engage in a life-or-death struggle against the Zangyack. One after the other, strong enemies gather to stand in the Gokaigers' way. Will the space pirates prevail over this brutal empire?