Jacks-of-all-trades Yun and Haberu investigate sounds emanating from an empty mansion. They locate the source nearby, while Mei hears the same tune.

Yun, Goro and their robot Jet Jaguar clash with a hostile pterosaur-like kaiju, Rodan. Mei travels to Tokyo to meet with a world-renowned researcher.

A legion of Rodan swarm towards Nigashio. The Otaki Factory crew scramble to lure them away from the city, but soon find themselves surrounded.

Following a group of Manda, a giant kaiju makes landfall in Tokyo. Red dust forces Mei and Professor Li's flight to London to divert to Milan.

Ground forces under the command of Matsubara respond to Godzilla's attack on Tokyo. Now in London, Mei attempts to decipher Ashihara's research.

Godzilla continues to advance on Tokyo, covering the city in red dust. The Otaki crew rushes to rescue a person ensnared by spider-like kaiju.

Mei learns of a large explosion triggered by Ashihara's calculator long ago. Kanoko gives Matsubara the Orthogonal Diagonalizer to use against Godzilla.