As the gang nears Neville and his men, their hopes of finding Danny grow until a shocking discovery threatens to impede their efforts. Meanwhile, orders from Monroe have upped the value of Danny's life and Nora connects with a fellow Rebel.

When Nora encounters a close relative, she must make a difficult choice between family and a promise. Meanwhile, one of the most ruthless members of the Militia is tracking Miles, Charlie and the gang and General Monroe hands Captain Neville a stern warning.

In the gang's quest to rescue Danny, the time has come for Miles and Monroe to meet face to face. Meanwhile, on the verge of reuniting with her brother, Charlie is conflicted by a shocking discovery and Rachel tests Monroe's patience.

Neville faces danger and hard choices as Monroe doubts his loyalty, he disowns his son Jason and gets captured by Miles. Rachel and Charlie begin slowly patching their relationship only to be pulled apart again as Rachel suddenly embarks on a quest to restore the power and her self-respect. She must make a dangerous journey to the mysterious Tower as it holds her only chance at redemption.

Life-threatening dangers abound on all sides as the Rebels infiltrate the Tower. Miles' leadership abilities and character are sorely tested as well as his feelings for Rachel and Nora. Aaron's genius comes in handy while Tom Neville and Randall Flynn bring new meaning to the dark side.

The relationship between Rachel and Gene remains strained as Charlie shows her mom how much she's grown. Meanwhile, Neville gives Jason a determined proclamation.

When Miles leads Monroe and Rachel south of the border; they're met with much more than they bargained for as Neville and Julia dance a delicate dance. Meanwhile, Charlie and Gene continue their search for Aaron.

Monroe finds himself in unfamiliar territory and must make a choice about his son Conner, while Neville and Julia continue to tango at Jason's dismay. Meanwhile, Gene leads Charlie to an uneasy discovery.

When Gene decides to assist the town of Willoughby, Rachel and Charlie feel compelled to help too as Miles and Monroe continue to join forces in order to survive.