Miles, Aaron and Rachel have found themselves in a mysterious small town, in the great nation of Texas where Rachel unexpectedly encounters an important figure from her past. Charlie finds herself on a mission in the Plains Nation while Neville and Jason search a refugee camp for a lost loved one. Meanwhile, Monroe has discovered a gritty role in his new environment.

Rachel makes a valiant attempt to rescue Miles and to escape with an ailing Tribesman's wife, while Neville and Jason are caught off-guard, assaulted by violent refugees. Charlie and Adam are at odds over what to do with Monroe. Legions of Tribesmen threaten all of Willoughby but with questionable leadership and Aaron faces death, yet again.

Aaron and Rachel speculate on the extreme effects of Nano technology, while he also confides in Cynthia. Meanwhile a figure from Miles's rugged past, Texas Ranger John Fry reemerges. Fry could be Miles' chance to bring down the Patriots in a big way, however Fry will need convincing. Charlie has a surprise for Miles and her mother, while Secretary Allenford holds a secret important to Neville.

When Rachel discovers some unsettling truths, Miles does what he does best to overcome obstacles before him and those he cares about. Meanwhile, Charlie shares a moment with her mom as Neville and Jason continue with their struggles between father and son.

The relationship between Rachel and Gene remains strained as Charlie shows her mom how much she's grown. Meanwhile, Neville gives Jason a determined proclamation.

Monroe finds himself in unfamiliar territory and must make a choice about his son Conner, while Neville and Julia continue to tango at Jason's dismay. Meanwhile, Gene leads Charlie to an uneasy discovery.

Monroe leads Connor and Charlie on a risky mission, as Miles and Rachel keep an eye on things in Willoughby. Meanwhile, Neville gets a new assignment and Aaron reunites with an old friend.

Aaron awakes to find himself in a world with power, married to Priscilla and running his company. As he gets the feeling that something isn't quite right, familiar faces begin to appear.

While Dr. Porter and Miles venture into Willoughby in search of allies within the town, Monroe seizes an opportunity to surprise the Patriots. Neville’s political maneuverings put him and Jason in serious danger, as Aaron and Priscilla continue on their journey.

As Miles, Monroe, Charlie and Connor rush to escape Austin, a confrontation with Texas Rangers separates Miles from the group. While out searching for Miles, Charlie is confronted by Neville who demands information on Jason's whereabouts. As a wounded Miles faces increasingly hopeless circumstances, Aaron learns that the nanotech has been busy.

United by their desire to take the Patriots down, Neville and Monroe form an uneasy alliance. As Rachel comes face-to-face with the nano-tech while searching for Aaron, things don't go according to plan for Miles, Charlie and Gene when they try to steal the Patriots' supply of mustard gas.