Congressional hopeful Sue Sylvester starts a smear campaign to ruin the candidacy of her biggest rival. Puck falls in love, and glee club directors Will and Shelby encourage some friendly competition between their clubs

Much to Blaine's chagrin, his big-shot Hollywood actor brother, Cooper, comes to visit.

Hopes and schemes play out as the Glee Club holds auditions for the school’s musical.

The kids of New Directions go head-to-head in their annual boys vs. girls mash-up competition, this time using the music of the movies. Meanwhile, in New York, Santana makes herself at home in the loft a little too quickly for Rachel and Kurt's comfort.

The glee club learns the dance of the hour: twerking. Meanwhile, Marley discovers Jake's relationship with Bree, and Rachel tries to convince Kurt they both should get tattoos.

Brittany and Santana plan their wedding. The Gleeks weekly assignment is the music of Burt Bacharach, and reigning diva Mercedes is recruited to act as a mentor for the new singers. Brittany learns a shocking family secret.