It's a new era at Glenbogle and Paul is the new laird. Despite his best interests, his attitudes towards the highland traditions are starting to bother the family. Meanwhile, Lexie returns briefly from New Zealand causing Paul to become a little suspicious.

Hector's brother Donald MacDonald returns after a 40-year absence to Glenbogle. But this surprise visit is not a welcome one for Molly who holds a mysterious grudge against Donald. Meanwhile, Lexie avoids the thought of returning to Archie in New Zealand by throwing a 60th birthday party for Molly.

It's the day after Paul and Lexie's kiss and while Paul is eager to discuss it, Lexie wants to forget it and decides to leave to New Zealand immediately. But slowly she begins to think that leaving her home that is Glenbogle might not be what she really wants. Then, the secret that is the cause of Molly's hatred towards Donald is revealed.

Ewan and Donald find a German Second World War fighter plane at the bottom of the loch and when Paul's former army officer Alex Faversham comes to investigate the plane, he digs up some unwanted memories in Glenbogle.

Alex has decided that the estate is not being run with a firm enough hand and takes it upon himself to bring his military organization to the Glenbogle household but it isn't long before everyone becomes annoyed with Alex. Meanwhile, while Molly is away, Golly strikes up a new friendship with a local dance teacher named Meg.

Isobel's farm is under threat when she can't pay the monthly rent on time, again and is persuaded by her old work colleague to return to Edinburgh to start up a new partnership. Meanwhile, a scruffy stranger who has lost his memory turns up on the estate and the family tries to help the hiker remember his past.

While Golly is away on holiday with Meg, Jess takes over as ghillie. Her skills are put to the test right away when poachers invade the estate. Meanwhile, Paul and Isabel join their new neighbor Lucy at a cocktail party and Paul and Lucy share an intimate moment spotted by Isabel.

Competition sparks between Paul and Chester when they both want to market Isabel's relish. To decide who will be the winner, a duel is called to see who is the better marksman of the two. Meanwhile Meg feels disheartened when Golly gets people into her dance class through favors and Jess gets set up on Rob Roy Radio.

Ewan is shocked when he finds his dad with another woman and learns that his parents are planning on separating. Meanwhile, Lucy's parents are coming over to meet Paul and begin to hint about the possibility of a future wedding.

The time has come for Paul's official coronation as laird of Glenbogle but the family is shocked to find out that Donald is the true heir to the title. Donald takes the news well and begins changing the estate in every which way including taxing the tenants. To settle the decision over who should be laird, Paul and Donald face off in a duel to prove themselves worthy. Meanwhile, Meg returns to Glenbogle with a shocking surprise of her own.

Ever since Meg passed away, Golly has been taking care of baby Cameron as a single parent, with the help of his daughter, Jess. Meanwhile, as spring returns to Glenbogle, the list of repairs to be done is endless and Paul finds himself face to face with a feisty young shepherdess named Iona.

Paul's goddaughter Amy shows up, a relentless troublemaker, and begins to shake up Ewan and Zoe's relationship. Then, Tam Cooper, a recently released convicted poacher returns to Glenbogle, creating havoc for Paul and Golly.

Paul launches his new idea of turning Glenbogle into a luxury retreat when a wealthy couple come to stay at the house. But when Golly has a back injury and Molly rushes to his aid, Paul finds himself short staffed and strapped for cash.

Ewan believes he's struck gold when he discovers that Amy has photographed a monster emerging from the loch. Although skeptical at first, Paul too soon finds himself captivated by the mystical creature and the magical power it seems to have.

Iona is stunned when her former fiancé, Malcolm McRae, comes to Glenbogle to help save the estate's forest from a spreading fungi. The surprise forces Paul to consider his true feelings for Iona while attempting to heal the damage in the woods. Meanwhile, Jess competes in the local Ghillie's challenge, as the only woman, with all odds stacked against her.

Emotions are running high. The bank can no longer wait for Glenbogle to pay off their debt, forcing Paul to sell the estate. Then, a wealthy stranger comes to buy the estate forcing the family to face their futures without Glenbogle. Meanwhile, Molly must decide between Golly and Killwillie, Donald must face up to his future, alone, Paul must confront his true feelings for Iona and Golly must confront his true feelings for Molly. All of this comes to a head, and a surprising visitor from the past lends a helping hand.