Jane looks for a way for Maura to meet her biological mother, Dr. Hope Davis. Angela gets involved in a political campaign. And a nun from Jane and Frankie Jr.’s past returns.

When a rich financier is found stuffed in a trashcan wearing a cloak and mask, Jane and Maura must figure out what happened. Meanwhile, Maura makes a deeper connection with her biological mother, Dr. Hope Davis, and half-sister, who still doesn’t know Maura is related to them.

Dr. Hope Martin returns to ask Maura for a life-altering favor. Jane worries about the impending spinal surgery for Lt. Col. Casey Jones (guest star Chris Vance), and Tommy shows off baby T.J. But all gets shoved to the side when a building collapses, trapping loved ones inside.

Jane is unhappy when a former colleague returns as the new lieutenant of the drug unit; a tip from Cailin causes Maura to investigate Hope.

On the first day of Paddy’s murder trial, Maura gains insight into her father from Hope. Korsak opens up about Cavanaugh’s tragic past. And in an unexpected twist, Maura must decide which side she’s on when she and Jane unravel a shocking secret about Paddy and Hope that affects them all.

When a food truck chef dies suddenly, all signs point to poisoning; Tommy announces some big plans; Maura's birth mother tries to make amends with her daughter.

When a wealthy woman is found dead at the bottom of her stairs, we realize that this was no ordinary slip-and-fall accident. The team is pressed into a game of cat and mouse with a clever killer determined to get away with the perfect crime. Plus, when Maura decides to undergo minor surgery, the surprising part of her healing will be reconnecting with Hope.