Daggett and Norbert learn how to mark territory with their scent glands. Their fraternal jealousy kicks off a marathon of territory claiming across the dam, the forest, and the entire country. The boys finally learn their lesson at the hands of some rather smelly lumberjacks.

While on their way to a local sports coliseum, Daggett annoys the bus driver who accidentally drives the bus through "a hole in the fabric of space and time." The boys end up at the real Roman Colosseum, where Daggett picks so many fights both beavers are forced to become gladiators. After surviving man-eating lions and the dreaded Two-Eyed Cyclops, the Beavers lead Caesar's army on a madcap chase that topples Rome.