Alto Saotome unexpectedly meets Ranka Lee and Sheryl Nome again at an observation park. However, the Vajra also shows up in the skies above the three, so they immediately head to the park's emergency shelter and get trapped inside.

Ranka Lee is contracted to act in a minor role during a movie about the legend of Shin Kudo, while Sheryl Nome is the main female protagonist of the film.

Ozma promises Ranka to attend her first live concert on Macross Frontier, but he is ordered to defend the fleet from another Vajra attack before that.

The Frontier populance's hatred of the Vajra is growing by leaps and bounds, and even Alto and Luka cannot control their own hate. Moreover, now armed with new prototype weapons, Alto and Luka consider using them to utterly exterminate all Vajra. However, Ozma (who has been researching Leon's conspiracy) is beginning to realise the identity of the real enemy behind the conflict. Meanwhile, Sheryl decides to sing again with a brand new perspective.