Nepgear, Uni, Rom, and Ram are instructed to go back home after their sisters' capture, but they can't bear to leave their families behind in danger. To get back their sisters, they'll have to learn how to transform into goddesses themselves.

Neptune's maybe having a tiny bit of trouble bouncing back, and Plutia's maybe getting a tiny bit irritated about it. Soon, though, Gamindustri finds itself facing a problem even bigger than Plutia's rage...