Girls that have been dubbed the "Progress" are those which have awakened to the superpower "Exceed". These girls, from the five different worlds that exist, are kept safe and trained at Seiran Academy. They train day and night with their alpha drivers, a person who has the ability to enhance their powers.

The girls have to deal with the aftermath of the Ouroboros attack. Their fighting power has been greatly reduced, and they soon find out that things are worse than they had feared when some of the members are corrupted by the darkness...

After saving the four other worlds through the portals, the defenses around Seiran island are strengthened in anticipation of a final attack. However, events do not proceed as anticipated...

In a final effort to save the worlds, Saya seeks to reconnect with Amane through a fault detected in the process used to imprison the alpha drivers.