Girls that have been dubbed the "Progress" are those which have awakened to the superpower "Exceed". These girls, from the five different worlds that exist, are kept safe and trained at Seiran Academy. They train day and night with their alpha drivers, a person who has the ability to enhance their powers.

The blooming battle between Saya and the other Progress comes to an end, and the result drives a wedge between Saya and Amane.

The girls have to deal with the aftermath of the Ouroboros attack. Their fighting power has been greatly reduced, and they soon find out that things are worse than they had feared when some of the members are corrupted by the darkness...

Saya and the others are heading to the Red World, Terra Rubiri Aurora, to check on the Ouroboros, but Elel's best friend, Ramiel, is waiting for them. Ramiel asks Elel if they're really friends.

In order to destroy the White World's world crystal by injecting a virus into it, Carene must fly at an incredibly fast speed. The only one who can stop her now is Stella and her Exceed with the power of super-speed. With the help of Saya and the others, Stella chases after Carene.

Nya figures she has to move after seeing Saya and the others deepen their bonds, and heads off to the Green World on her own. But an army of Ouroboros stands in her way.

After saving the four other worlds through the portals, the defenses around Seiran island are strengthened in anticipation of a final attack. However, events do not proceed as anticipated...

In a final effort to save the worlds, Saya seeks to reconnect with Amane through a fault detected in the process used to imprison the alpha drivers.