When the evil mayor of Oakton decides to bulldoze Liberty Park and build a dangerous amusement park in its place, Surly Squirrel and his ragtag group of animal friends need to band together to save their home, defeat the mayor, and take back the park.

Surly, a curmudgeon, independent squirrel is banished from his park and forced to survive in the city. Lucky for him, he stumbles on the one thing that may be able to save his life, and the rest of park community, as they gear up for winter - Maury's Nut Store.

After breaking out of a rehabilitation zoo, Frankie, a squirrel with all the angles, returns to his forest. He learns from his dim-witted best friend, Cody, a porcupine and street-con, that while he was in lock-up, the entire forest was picked clean by The A.C.O.R.N.S. Institute (Alternative Cosmetic Oils for Regenerative Natural Skin-care). No acorn was left behind. With winter just around the corner, it's a potentially devastating blow to all forest critters. So, Frankie has no choice but to bust into the brand new, state-of-the-art, rodent-proof A.C.O.R.N.S. Processing Plant and take back all of the acorns! Though a seemingly impossible heist, success would not only save the lives of his forest friends, but it would also give Frankie a chance to win back the love of his life, Lola, a local "show squirrel." Sadly, Lola has been burnt by too many of Frankie's false promises in the past.

When Petra spends an evening with her best friend Sanna, Sanna asks how she knew she was bi. Petra tries in earnest to show her the truth.