Based on the book O Fantástico Mistério de Feiurinha, the story combines real world and fantasy, dream and determination, with a surprising end, even for a fairy tale. The movie shows the fight of the fairy tale characters, now adults, who come together to recover the fable of the Little Ugly Princess, a beautiful Princess with a strange name, which only makes sense for those who know her story. The problem is that children and adults have forgotten the story of the Little Ugly Princess, which is as wonderful as other major fables for children. Cinderella brings together other fairy tale characters to meet the writer who keeps, lost in a small notebook, the Little Ugly Princess story.

O que acontece depois do "viveram felizes para sempre"? As princesas e os príncipes têm filhos, probleminhas com as sogras, problemas nos relacionamentos, engordam, mas... continuam felizes. Entretanto, prestes a fazerem bodas de prata, as princesas Cinderela, Rapunzel, Branca de Neve, Bela Adormecida, Bela da Fera, com a ajuda de Chapeuzinho Vermelho, precisam desvendar o mistério do desaparecimento da princesa feiurinha. Caso contrário, todo o reino encantado poderia desaparecer.