Omega Chi and Kappa Tau each throw parties to celebrate the end of the campus restrictions.

The students enjoy spring break on the beach. Cappie and Casey explore their new friendship at a time when Rebecca needs Cappie's attention. Evan looks to move on and escape his feelings and ends up with a familiar person. Rusty returns to school and on his way back he reunites with old friends.

Casey and Ashleigh discover that they have the same crush for the ZBZ Crush Party. Meanwhile, Rusty meets his new RA and turns to him for help.

Casey and Rusty organize a casino night in order to help a debt-plagued Ashleigh. Elsewhere, Evan struggles with accepting the fine print on his trust fund. Cappie's in need of new car. And, Rebecca wants more from a relationship but she's not sure if she'll get it from Cappie.

It's the ZBZ Formal and Casey and Max decide to go together. At the event, Ashleigh and Rusty bond. Also, Rebecca decides to skip the event claiming she needs a break from sorority events.

Frannie and Casey sling mud at each other as the sorority presidential election approaches; Calvin faces the consequences after he cheats on Michael.

Rusty finds out a secret about Dale's high school friend and fellow Purity Pledge member, but is uncertain to tell Dale what he knows. Meanwhile, Ashleigh, as the new ZBZ president, finds herself in the middle of a battle between Casey and Frannie as they pursue the Omega Chi's for partnership for the coming school year.

Casey faces a hard decision when she learns that Max is planning to go to Cal Tech for grad school and she is offered an internship. She turns to Lauren Conrad (who appears in a dream sequence) for advice. Also, the pledges are put through Hell Week and Rusty is driven to face the challenge no matter what he is forced to do.

Now a sophomore, Rusty gets to participate in Rush Week for the first time and he is determined to land Andy as a pledge for Kappa Tau, but Calvin feels the pressure from Evan to get Andy to pledge Omega Chi. Who will prevail in the battle for the big man on campus? Ashleigh and Casey start to doubt Rebecca's "mole" abilities and begin to question whether she's actually a double-agent for Frannie's new sorority when the ZBZ girls’ rush week “Project Runway” fashion skit is successfully sabotaged by Frannie.

When Casey discovers that Cappie is taking the same women’s studies class taught by Professor Freeman, she knows that it will complicate her relationship with Max. As a result, Casey decides to enlist the help of her new professor to help Max and Cappie get along. Meanwhile, Rusty starts his first elective class in Art History, and as a lucky bonus meets a pretty new transfer student, Jordan. Although he’s nervous about talking to her, he finds the courage to invite her to a KT party. But the night of the party he learns that some guys have all the luck, namely the new KT pledge, Andy. At the ZBZ house, Ashleigh hires a new hasher, Fisher, who draws a lot of attention from the sisters of ZBZ but, Fisher only has eyes for Ashleigh.

A lonely Casey meddles in Rusty's new romance with Jordan. Elsewhere, Calvin finally gets a new roommate, and then he discovers that the guy has a surprising secret; and Evan and Cappie reignite their rivalry, this time over a secret-society member.

Rusty and Ashleigh try hard to win the float competition, while Casey and Cappie try avoid each other as Homecoming Week heats up. Elsewhere, Evan gets word that he may lose his trust fund, and Ashleigh uncovers one of Fisher's secrets.

The students at CRU feel the ramifications of the Kappa Tau's "End of the World" Party. Rusty risks being dropped from the honors program if he cannot raise his grade in organic chemistry and Casey adjusts to single life.

Casey finds out about Rebecca and Fischer's secret kiss, but when Rebecca confesses to Ashleigh, she outs Casey for keeping it a secret. When Mr. Cartwright, Mr. Howard, and Mr. Logan arrive for Father-Daughter weekend at ZBZ, they find out about their daughters' feud, and it reflects upon their weekend behavior. At the same event, Rusty gets nervous about meeting Jordan's father. Meanwhile, in an effort to get over their lost loves, Dale and Cappie fawn over the same girl, but they both ultimately get rejected.

It's time for the annual all-Greek "Undie Run", and the Pan-Hellenic board voted strongly against it. In an effort to change the vote, Casey uses her political savvy to enlist IFC president, Evan, to stage a petition signing. Meanwhile, it's nearing the end of the Greek-wide game of Gotcha!, and only Rusty and Jordan are left, which puts a damper in their effort to spend alone time together. Additionally, Evan finds out about Calvin and Grant's relationship and helps them keep it a secret from the rest of Omega Chi.

Shocked to learn that ZBZ has slipped in the ranks down to fourth, Casey is determined to bring ZBZ back to the number one spot by motivating their pledge class to take action, believing that, if they create a cool photo shoot for the Lamda Sig calendar, it could improve their house’s ranking. However new pldge Abby convinces the other pledges to boycott the shoot so, in order to make nice with the ZBZ pledges, Ashley and Casey get them all makeovers. Meanwhile, dissatisfied with her college experience, Jordan wonders how to make her time at CRU more fulfilling and, in an effort to prove his love and commitment for Jordan Rusty decides to lavaliere her. When Rusty divulges his plans to his KT brothers, they warn him of the Kappa Tau curse that occurs when a brother lavalieres his girlfriend. Elsewhere, Cappie helps Evan find a part time job to help his financial situation, but when he meets one of Evan’s new and very attractive co-workers, Lana, he decides to get a job too.

Everyone is packing their bags and heading home for Thanksgiving, but Casey and Rusty find themselves stranded at CRU when their parents leave them behind for Maui. Will Thanksgiving at CRU beat going home for the holidays? A surprise at the end will leave you speechless and wanting more.

Cappie and Casey go public, and Omega Chi and Kappa Tau get serious about pranking. Things heat up when Song Fest pits ZBZ against Gamma Psi. Rusty and Dale take their rivalry out of the lab.

The Cyprus Rhodes students are preparing for the "Help Us Rebuild Our Castle Fairytale Ball" for the Gamma Psi's - some out of guilt - by picking out their best prince and princess attire.

The ZBZ's are still on edge when they hear their candle is responsible for the Gamma Psi fire, and Casey cracks under the guilt, putting the ZBZ house in jeopardy! Hoping to boost morale and her marketing major, Ashleigh enlists Rusty and Dale in a philanthropic "Geek Auction," but it's up to Casey to play dirty to save the ZBZ's, their house and their Greek rank. Find out if Casey can save the day.

The KTs and OXs are preparing for the Gods of Golf Tournament, the winner of which will be named the top fraternity on campus. Meanwhile, the ZBZs are disappointed to learn that the annual Greek Week has been canceled. Casey talks to Katherine, who agrees to bring back the Greek Goddesses Pageant, under the condition that the competition focus more on strength of character and less on looks, of course. Which houses will end the year on top?

The season opens with Casey, Evan, & Ashleigh’s graduation day. Casey and Cappie haven’t spoken since Spring Break and are anticipating saying their final goodbyes to each other, which does not go as smoothly as expected. We then skip to the end of summer and Casey packing up to move to Washington DC for law school. Through some prompting from Evan, Casey learns that the reason she did not get into CRU law school was a result of a bad recommendation from Joel (Sam Page). After Casey confronts Sam, he reconsiders and has his letter rescinded. Casey is accepted into CRU law and decides to stay after all. Ashleigh has moved to New York for an internship that is not exactly what she was hoping for. Evan is attending CRU law school with Casey.

The KT’s decide since they lost several members of their frat last year, it’s time to actually try to recruit pledges. The KTs set up a booth during rush on campus to help with recruiting. Tegan (Charisma Carpenter) shows up to help the ZBZs maintain their newly regained best house on campus title. Tegan and Rebecca argue on how rushing should go and Casey comes to Rebecca’s rescue with a plan to have Dana (Martha MacIsaac) keep Tegan distracted. The ZBZs entertain a potential new pledge Heather, who is Heath’s sister. Dale also entertains the idea of rushing this semester

Rusty, Dale and Dana are hanging out together and Dana suggests throwing a murder mystery dinner party to help Dale socialize. Members of Omega Chi Delta then storm in and take Dale away as he‘s pledged to them. Casey gets a surprise visit one night from Ashleigh, who has lost her job in New York and now plans to stay back at the house. Calvin reveals that he has no major, just like Cappie, but wants to keep it a secret. Meanwhile, Rusty worries that Dale is joining Omega Chi to get back at him, so he devises a plan to get Dale membership into Kappa Tau. But Dale is insulted by Rusty‘s insistence, and goes to Omega Chi‘s hazing anyway. While he‘s there, Rusty and his friends kidnap him and take him to Dana‘s murder mystery party, inadvertently leaving Spidey there to face the Omegas‘ wrath. Dale eventually leaves Rusty‘s party, claiming that he wants to be a part of that fraternity, and that Rusty should care about his own pledge Spidey instead. Casey is having a hard time finding a study group to join in her new law class because of her controversial past. Ashleigh reveals that she actually ran away from her job because she couldn‘t handle it. Katherine (Nora Kirkpatrick) returns to attend CRU Law school.

Kappa Tau faces academic probation when one of their pledges mysteriously receives straight F‘s, the KT‘s suspect Omega Chi and Dale are behind it. Rebecca uses the investigation as her post Evan break up coping mechanism and discovers that Omega Chi is actually playing a trick on Dale. Evan takes revenge on Casey for his break up with Rebecca by attempting to make her look less than capable in her Law school studies.

In the series finale, the Kappa Taus try to stop the demolition of their house. Meanwhile, Cappie and Casey come to a realization about their on-again, off-again romance; and Cappie and Evan determine whether their friendship is meant to last.