Omega Chi becomes the victim of Kappa Tau's pranks when Cappie seeks to get even with Evan for blowing the whistle on their recent underground festivities. Meanwhile, Casey's rivalry with Rebecca intensifies when she learns about Rebecca's new relationship.

Omega Chi and Kappa Tau each throw parties to celebrate the end of the campus restrictions.

Spring Break is almost here, Cappie gets his pledges to practice with their fake ID's, leading Rusty into a tricky situation. Meanwhile, Casey goes to Evan for help with her LSAT test and in the process she begins to question her future.

The students enjoy spring break on the beach. Cappie and Casey explore their new friendship at a time when Rebecca needs Cappie's attention. Evan looks to move on and escape his feelings and ends up with a familiar person. Rusty returns to school and on his way back he reunites with old friends.

Casey and Rusty organize a casino night in order to help a debt-plagued Ashleigh. Elsewhere, Evan struggles with accepting the fine print on his trust fund. Cappie's in need of new car. And, Rebecca wants more from a relationship but she's not sure if she'll get it from Cappie.

Rusty is looking into the list of Kappa Tau actives and this makes him very nervous about one member's true identity being uncovered. Also, Casey seems to like Max, Rusty's R.A. And Ashleigh accepts to accompany Calvin and Michael to a gay bar as Michael says Calvin doesn't have enough gay friends.

Tired of the poor treatment they've received the pledges at Kappa Tau fight back. Elsewhere, Evan conspires to use Calvin and Michael's relationship to help a brother who's failing.

The ZBZ women attend their national sorority convention; a party for the engineering department gets out of control; Evan takes advantage of Frannie's absence.

It's the ZBZ Formal and Casey and Max decide to go together. At the event, Ashleigh and Rusty bond. Also, Rebecca decides to skip the event claiming she needs a break from sorority events.

Casey faces a hard decision when she learns that Max is planning to go to Cal Tech for grad school and she is offered an internship. She turns to Lauren Conrad (who appears in a dream sequence) for advice. Also, the pledges are put through Hell Week and Rusty is driven to face the challenge no matter what he is forced to do.

Now a sophomore, Rusty gets to participate in Rush Week for the first time and he is determined to land Andy as a pledge for Kappa Tau, but Calvin feels the pressure from Evan to get Andy to pledge Omega Chi. Who will prevail in the battle for the big man on campus? Ashleigh and Casey start to doubt Rebecca's "mole" abilities and begin to question whether she's actually a double-agent for Frannie's new sorority when the ZBZ girls’ rush week “Project Runway” fashion skit is successfully sabotaged by Frannie.

When Casey discovers that Cappie is taking the same women’s studies class taught by Professor Freeman, she knows that it will complicate her relationship with Max. As a result, Casey decides to enlist the help of her new professor to help Max and Cappie get along. Meanwhile, Rusty starts his first elective class in Art History, and as a lucky bonus meets a pretty new transfer student, Jordan. Although he’s nervous about talking to her, he finds the courage to invite her to a KT party. But the night of the party he learns that some guys have all the luck, namely the new KT pledge, Andy. At the ZBZ house, Ashleigh hires a new hasher, Fisher, who draws a lot of attention from the sisters of ZBZ but, Fisher only has eyes for Ashleigh.

Troubled by the recent discovery that Andy hooked up with Jordan, Rusty is hesitant to take Andy on as his little brother. Will Rusty be able to get past the budding relationship to become the big brother Cappie has been to him? When Cappie hears of Rusty’s reluctance, he takes Rusty aside to remind him that he brought Andy as a pledge to the fraternity and it is his responsibility to be the best Big Brother that Rusty can be for him. Meanwhile, when Evan’s real-life brother, Patrick, comes for a visit, their celebration at Dobler’s is interrupted by Cappie and the three end up reminiscing about their old camp days. But later Patrick shocks and disappoints Evan by asking for money. Frustrated by his brother’s actions, Evan takes it out on Frannie. When Casey and Rebecca believe that the thief responsible for stealing the ZBZ food supply is Fisher, they can’t figure out why Ashleigh won’t accept it. But when they dig a little further, Ashleigh reveals the delicious truth.

Rusty attempts to solve a mystery involving a secret service he ran across on campus in hopes of impressing Cappie. Meanwhile, some of Frannie's defectors from ZBZ house have asked to return, and Rebecca seeks dating advice from Calvin as she readies to go out with Robin.

Rusty and Ashleigh try hard to win the float competition, while Casey and Cappie try avoid each other as Homecoming Week heats up. Elsewhere, Evan gets word that he may lose his trust fund, and Ashleigh uncovers one of Fisher's secrets.

After she falls into a manhole risking her life, coupled with the Kappa Tau's throwing an "End of the World" party, Casey is sent into an emotional tailspin, questioning her relationship with Max and her lingering feelings for Cappie. Meanwhile, Ashleigh faces her own punishment with Pan Hellenic after a complaint is lodged by Frannie for the Homecoming Float sabotage. Ashleigh turns to Casey to help fight the charge, but when Casey gets caught up in her own personal drama, she leaves Ashleigh to face Frannie alone. And with Dale having trouble keeping his cougar girlfriend's sexual advances at bay and Calvin trying his best to resist hooking up with his new hot roommate, Grant. Dale convinces Calvin to go in on a "purity pledge" partnership so they can help each resist their most recent temptations.

The students at CRU feel the ramifications of the Kappa Tau's "End of the World" Party. Rusty risks being dropped from the honors program if he cannot raise his grade in organic chemistry and Casey adjusts to single life.

Casey finds out about Rebecca and Fischer's secret kiss, but when Rebecca confesses to Ashleigh, she outs Casey for keeping it a secret. When Mr. Cartwright, Mr. Howard, and Mr. Logan arrive for Father-Daughter weekend at ZBZ, they find out about their daughters' feud, and it reflects upon their weekend behavior. At the same event, Rusty gets nervous about meeting Jordan's father. Meanwhile, in an effort to get over their lost loves, Dale and Cappie fawn over the same girl, but they both ultimately get rejected.

Shocked to learn that ZBZ has slipped in the ranks down to fourth, Casey is determined to bring ZBZ back to the number one spot by motivating their pledge class to take action, believing that, if they create a cool photo shoot for the Lamda Sig calendar, it could improve their house’s ranking. However new pldge Abby convinces the other pledges to boycott the shoot so, in order to make nice with the ZBZ pledges, Ashley and Casey get them all makeovers. Meanwhile, dissatisfied with her college experience, Jordan wonders how to make her time at CRU more fulfilling and, in an effort to prove his love and commitment for Jordan Rusty decides to lavaliere her. When Rusty divulges his plans to his KT brothers, they warn him of the Kappa Tau curse that occurs when a brother lavalieres his girlfriend. Elsewhere, Cappie helps Evan find a part time job to help his financial situation, but when he meets one of Evan’s new and very attractive co-workers, Lana, he decides to get a job too.

Casey is ready to discuss the future with a resistant Cappie, and the situation isn't helped by handsome Washington politician Joel (Sam Page), who encourages Casey to envision a potential career in politics after she volunteers to help him with his voter registration campaign at the Cyprus Rhodes campus. Find out if Cappie is ready to grow up or if he will fall behind.

It's Valentine's Day at CRU and love is in the air! Casey and Cappie are at war over who can come up with the most romantic Valentine's Day gift. Casey also tries her hand at playing Cupid when she sets Katherine up with Rusty in an attempt to get back in Katherine's good graces. Will Katherine and Rusty spark a love connection or will their awkwardness get in the way?

The KTs and OXs are preparing for the Gods of Golf Tournament, the winner of which will be named the top fraternity on campus. Meanwhile, the ZBZs are disappointed to learn that the annual Greek Week has been canceled. Casey talks to Katherine, who agrees to bring back the Greek Goddesses Pageant, under the condition that the competition focus more on strength of character and less on looks, of course. Which houses will end the year on top?

The season opens with Casey, Evan, & Ashleigh’s graduation day. Casey and Cappie haven’t spoken since Spring Break and are anticipating saying their final goodbyes to each other, which does not go as smoothly as expected. We then skip to the end of summer and Casey packing up to move to Washington DC for law school. Through some prompting from Evan, Casey learns that the reason she did not get into CRU law school was a result of a bad recommendation from Joel (Sam Page). After Casey confronts Sam, he reconsiders and has his letter rescinded. Casey is accepted into CRU law and decides to stay after all. Ashleigh has moved to New York for an internship that is not exactly what she was hoping for. Evan is attending CRU law school with Casey.

In the series finale, the Kappa Taus try to stop the demolition of their house. Meanwhile, Cappie and Casey come to a realization about their on-again, off-again romance; and Cappie and Evan determine whether their friendship is meant to last.