Emmitt Smith is an NFL Hall of Fame running back who, since retirement, has devoted himself to business and giving back to the community. But recently, he decided to embark on a journey inward, tracing his family's past. It was a journey that took him beyond his wildest expectations, discovering not only his African roots, but his connections to slave owners in Virginia.

Lisa Kudrow journeys to the small Jewish town of Ilya where she discovers how her great-grandmother died in the Holocaust. Yet, Lisa finds hope when she uncovers a 60-year old family mystery.

On his journey to learn more about his paternal roots, Matthew Broderick discovers the heroic story of his grandfather during World War I and finds the final resting place of his great-great-grandfather who fought in the Civil War.

Brooke learns a tragic secret about her grandmother and then discovers a connection to royalty through her paternal roots.

Susan leads an emotionally charged search for her family roots. Starting with her estranged grandmother Anita who abandoned Susan's mother in early childhood, Susan's journey takes her all the way to Tuscany, Italy. Back in America, Susan discovers a surprising connection to her grandmother.

Spike begins his journey in Atlanta, Georgia, where his grandmother Zimmie worked in the segregated Southern systems for many years. Here, Spike follows the trail of his ancestors dating back to the Civil War and discovers a shocking family connection.