Kuzco is a self-centered emperor who summons Pacha from a village and to tell him that his home will be destroyed to make room for Kuzco's new summer home. Kuzco's advisor, Yzma, tries to poison Kuzco and accidentally turns him into a llama, who accidentally ends up in Pacha's village. Pacha offers to help Kuzco if he doesn't destroy his house, and so they form an unlikely partnership.
Shaun bluffs his dimwitted farmer master into bidding for three llamas at a county fair. Once they arrive, however, they cause such chaotic destructive mayhem that Shaun has to carefully remove them – high-speed chases, careful rooftop scrambles and dangerous falls ensue.
A strict father-son relationship is put to the test when an unexpected surprise is brought into the mix.