Betty is assigned to interview a novelist. Amanda and Marc may have uncovered who Amanda's biological father is. Daniel unknowingly dates Wilhelmina's sister.

Betty plans to spend her 24th birthday accompanied by Henry, but Charlie's sudden appearance jeopardizes her plans. Meanwhile, Wilhelmina plots against Renee and Daniel's relationship by hinting at secrets from a hidden past in her sister's life.

Renee becomes convinced that Betty is in love with Daniel and thus starts a rivalry between the two for his affections. In the meantime, Gina returns to Queens to flash her new life and rich husband before Hilda's face.