Rebecca makes a surprise visit to Randall's home where she encounters his biological father, William. Kevin anxiously decides to take the plunge and move to New York to pursue a Broadway job, with the expectation that his sister, Kate, will move with him. Toby is determined to get Kate out of Kevin's shadow. Jack and Rebecca take home their babies from the hospital and adjust to the new life of parenthood, while coping with the loss of their other child. Rebecca struggles to connect with their adopted child and searches for answers.

Kate, Kevin and Randall head to their family's cabin to get away after their chaotic family Thanksgiving. A very angry Randall finds comfort and explanation from his father, Jack. Olivia speaks a harsh truth to Kate, which drives a wedge between Kate and Kevin. Rebecca and Jack struggle with what to do when 9-year-old Randall tells them that he's been looking for his birth parents.

As the birth date of her triplets quickly approaches, a frazzled and emotional Rebecca forgets Jack's birthday. Dr. Katowski struggles with the loss of his wife and wonders how he'll be able to continue on with his life without her. Joe, the firefighter who delivered baby Randall to the hospital, fights to save his marriage.

Randall and William take a road trip to Memphis, where Randall learns about his biological father's past.

It's the Big Three's 37th birthdays. Randall and Beth debate a big life change, Kate takes the first step in pursuing a new passion, and Kevin balances the demands of his career and relationship. Meanwhile, Jack and Rebecca deal with the fallout of their big fight.

Randall adjusts to the foster system. Kate and Toby take the next step in their relationship. Kevin visits Sophie in New York. Jack and Rebecca finalize Randall's adoption.

Randall and Beth are faced with a hard choice. Jack takes Randall on a college tour.

Deja's life with her mom, Shauna, and great grandmother Joyce unfolds from the very beginning. Randall and Beth welcome visitors to their home.

The Pearson family gathers for their first Thanksgiving at Randall's Philadelphia townhouse.

The Pearsons gather to celebrate their 40th birthday while musing about the day they were born.

Kevin visits Randall in Philadelphia.