La historia de Ikoku Meiro No Croisée toma lugar en la segunda mitad del siglo 19, donde la cultura japonesa gana popularidad en Occidende. Es ahí donde conocemos a una joven japonesa de nombre Yune, que acompaña a un viajero fránces llamado Oscar en su viaje de regreso a Francia ofreciendose a ayudar en el taller de herrería de su familia en París: Los sobrinos de Oscar y Claude la dueña de la tienda aceptan a regañadientes cuidar de Yune, confome transcurre la historia nos vamos enterando como estos dos personajes que tienen tan poco en común llegan a entenderse y convivir en París en los inicios de 1800.

Yune eats her first French breakfast and attempts to hide her dislike of cheese! Claude, Oscar's grandson, takes Yune shopping.

Claude becomes inspired by Yune with an idea for a sign for a nearby music shop. Later, they find the owners of Yune's kimono.

Alice, the youngest of the upper-class Blanche family, becomes interested in Yune and wants her to stay with her, but Yune has made a promise to the Claudels.

Claude scolds Yune for her kindness, which is taken advantage of by a boy who steals from the shop.

Alice pays a visit to Yune at Oscar's shop and invites her over for a photograph. There, Yune learns of a past between Alice's sister Camille and Claude

Yune faints from a fever, which fills Claude with guilt for scolding her earlier. He receives help from Alice and her doctor to cure Yune.

A tea party at the Blanche's brings Claude and Camille together, while Alice shows Yune the nursery where she grew up.

Claude speaks about the past with Camille when they used to play together in the Blanche's family garden.

Oscar spends time with Yune and Alice. He shows off a slide projector that reminds Alice of her parts of her life back in Japan.

Yune goes out to search for a lost black cat and gets lost, which worries Claude who cannot find her.